Journal Articles

2022 Moorad, J.A. and S. Ravindran. Natural selection and the evolution of asynchronous agingThe American Naturalist. 199(4)

2020 Ivimey-Cook, E.R. and J.A. Moorad. The diversity of maternal age effects upon pre-adult survival across animal species. Proceedings of the Royal Society B:20200972.

2020 Ratz, T, A Perodaskalaki, J Moorad, and P Smiseth. Effects of inbreeding on behavioural plasticity of parent-offspring interactions in a burying beetleJournal of Evolutionary Biology. 33(7):1006-1016.

2020 Moorad, JA, DEL Promislow, and J Silvertown. Williams' intuition about extrinsic mortality is irrelevantTrends in Ecology and Evolution. 35(5):379.

2020 Moorad, JA, DEL Promislow, and J Silvertown. George C. Williams' problematic model of selection and senescence: Time to move onTrends in Ecology and Evolution. 35(4):303-305.

2019 Moorad, JA, DEL Promislow, and J Silvertown. Evolution of senescence: a reassessment of Williams' extrinsic mortality hypothesisTrends in Ecology and Evolution. 34(6):519-530*.2018 

2018  Ivimey-Cook, E.R. and J.A. Moorad. Disentangling pre- and post-natal maternal age effects on offspring performance in an insect with elaborate maternal care. American Naturalist. 192(5):564-576.

2018  Corbett, S, A Courtiol, V Lummaa, J Moorad, and S Stearns. The transition to modernity and chronic disease: mismatch and natural selection. Nature Reviews Genetics. 19(7):419-430.2017

Moorad, JA and CA Walling. Measuring selection for genes that promote late-life human lifespan. Nature: Ecology & Evolution. 1:1773-1781.  Access the recommendation on F1000Prime

2016 Moorad JA and DH Nussey. Evolution of maternal effect senescence. Proceedings of the National Academy of Sciences 113(2):362-367.

2015 Hayward, AD, JA Moorad, CE Regan, C Berenos, JG Pilkington, JM Pemberton, and DH Nussey. Asynchrony of senescence among phenotypic traits in a wild mammal population. Experimental Gerontology 71:56-68*.

2015 Pilakouta, N, S Jamieson, JA Moorad, and PT Smiseth. Parental care buffers against inbreeding depression in burying beetles. Proceedings of the National Academy of Sciences 112(26):8031-8035.

2014 Moorad JA. Individual fitness and phenotypic selection in age-structured populations with constant growth rates. Ecology 95:1087-1095.

2013 Moorad JA. A demographic transition altered the strength of selection for fitness and age-specific survival and fertility in a 19th century American population. Evolution 67(6):1622-1634.

2013 Moorad JA. Multi-level sexual selection: Individual and family-level selection for mating success in a recent human population. Evolution 67(6):1635-1648*.

2013 Moorad JA and MJ Wade. Selection gradients, the opportunity for selection, and the coefficient of determination. The American Naturalist 181(3):291-300.

2012 Moorad JA, DEL Promislow, N. Flesness, and RA Miller. 2012. A comparative assessment of univariate longevity measures using zoological animal records. Aging Cell 11:940-948.

2011 Moorad JA, DEL Promislow, KR Smith, and MJ Wade. Mating system change reduces the strength of sexual selection in an American frontier population of the 19th century. Evolution & Human Behavior 32:147-155.

2011 Moorad JA and DEL Promislow. Evolutionary demography and quantitative genetics: Age-specific survival as a threshold trait. Proceedings of the Royal Society B 278:144-151.

2010 Moorad JA and DEL Promislow. Evolution: Aging up a tree? Current Biology 20:406-409*.

2009 Moorad JA and DW Hall. Age-dependent mutational effects curtail the evolution of senescence by antagonistic pleiotropy. Journal of Evolutionary Biology 22:2409-2419.

2009 Moorad JA and DW Hall. Mutation accumulation, soft selection, and the middle-class neighborhood. Genetics 182:1387-1390.

2009 Moorad JA and DEL Promislow. What can genetic variation tell us about the evolution of senescence? Proceedings of the Royal Society B 276:2271-2278.

2008 Moorad JA and DEL Promislow. A theory of age-dependent mutation and senescence. Genetics 179:2061-2073.

2008 Moorad JA and TA Linksvayer. Levels of selection on threshold characters. Genetics 179:899-905.

2005 Moorad JA and MJ Wade. A genetic interpretation of the variation in inbreeding depression. Genetics 170:1373-1384.

2001 Philippi TE, MA Simovich, ET Bauder, and JA Moorad. Habitat ephemerality and hatching fractions of a diapausing anostracan (Crustacea: Branchiopoda). Israel Journal of Zoology 47:387-395.

2001 Moorad JA, MA Simovich, and MS Mayer. Identification of southern California branchinectid cysts (Crustacea, Anostraca) using RAPD-PCR species-specific markers. Transactions of the Western Section of the Wildlife Society 37:16-21.

1997 Moorad JA, MS Mayer, and MA Simovich. Extraction of DNA from anostracan cysts (Crustacea, branchiopoda) for use in RAPD-PCR analyses. Hydrobiologia 359:159-162.

* Invited paper

Invited book chapters

2016  Arbuthnott, D, DEL Promislow, and JA Moorad. Evolutionary Theory and Aging. Handbook of Theories of Aging. 3rd Edition. B. Kennedy, ed., Springer Publishing.

2016  Moorad JA and DEL Promislow. Evolution of aging and menopause. How Evolution Shapes Our Lives: Essays on Biology and Society. Lenski, R and J. Losos, ed., Princeton University Press.

2016  Moorad JA. Why Do We Age? Encyclopedia of Evolutionary Biology. Vol. 1. R. Kliman, ed., Oxford: Academic Press.

2013  Moorad JA and DEL Promislow. Evolution of aging and menopause. The Princeton Guide to Evolution. J. Losos, ed., Princeton University Press.

Invited book reviews

2016 Moorad JA. Review of Sociality, Hierarchy, Health: Comparative Biodemography: A Collection of Papers, edited by M Weinstein and MA Lane, Evolution, Medicine, and Public Health. 2016:67-68.


Nussey, D and J Moorad. The Evolution and Ecology of Senescence,  Oxford University Press (Under Contract)